AREADNE 2022 Logo   AREADNE 2022
Research in Encoding And Decoding of
Neural Ensembles

Nomikos Conference Centre
Santorini, Greece
28 June – 2 July 2022
  AREADNE Brain Labyrinth Logo


The Organizing Committee and Co-Chairs would like to thank each of the participants in AREADNE 2022 for their contributions to a highly successful meeting that promoted cutting-edge systems neuroscience from scales that ranged from the the tiny to the vast, from the activity of individual neurons in the fly to the planned mapping of the entire human brain. This part of the web site now holds the archival material from the conference.

The final scientific program is available as a PDF download from

Please cite the program as: N. G. Hatsopoulos, J. S. Pezaris (editors) Proceedings of AREADNE 2022, Santorini, Greece, 28 June – 2 July 2022, published by The AREADNE Foundation, Inc., Cambridge, Massachusetts, USA,

Information on past (and future) AREADNE conferences can be found at

Site Overview

Introduction (scroll down)
Important Dates
Call for Abstracts
Poster Information
Travel Information
Hotel Information
Practical Information
Restaurant Information
Additional Activities
Frequently Asked Questions
Scholarships and Resources for Travel Grants
Archives of Previous Meetings
Google Group for Official Announcements
Contact the Conference Organizers


A fundamental challenge in neuroscience is to understand how the activation of ensembles of neurons gives rise to the higher order functions of the brain including learning, memory, cognition, perception, action and ultimately conscious awareness. Over the past fifty years, electrophysiological recordings in behaving animals have revealed considerable information regarding what the firing patterns of single neurons encode in isolation, yet how collections of neurons interact to perform these functions largely remains a mystery.

Cactus and Caldera Recent technological advances have provided glimpses into the global functioning of the brain. These methods include multi-microelectrode array electrophysiology, optical imaging and stimulation, connectomics, functional magnetic resonance imaging, and genetic manipulation. Such advances have expanded our knowledge of brain functioning beyond the single neuron level into the realm of neural ensembles, networks of neurons whose coordinated activity represent and encode aspects of perception, thought, and action.

Gaining such insight on how neuronal ensembles carry information has enabled the development of brain-machine interfaces (BMI) to enhance the capabilities of patients with sensory or motor deficits. Particularly, research in how neuronal ensembles encode sensory stimuli has made it possible to develop perceptual BMIs for the hearing and visually impaired. Analogously, research in how neuronal ensembles decode motor intentions has resulted in motor BMIs, by which people with severe motor disabilities gain the ability to control external devices.

Conference Mission Statement

AREADNE Conferences have three major goals. First, to create a common touch-point for our widely disparate and hybrid field, by gathering global scientific leaders who work on neural ensembles. Second, to foster discussion and interaction between attendees and encourage the establishment of lasting professional relationships, through careful conference organization and planning. Third, to promote systems neuroscience in Greece, by creating a world-class forum for cutting-edge research.

Conference Location and Organization

Nomikos Entrance The 2022 conference will take place on the Greek island of Santorini from Tuesday through Saturday, 28 June - 2 July 2022, starting with an informal Welcome Reception the evening of Tuesday, 28 June 2022 -- this schedule is a day earlier in the week than previous meetings. The conference will be at the Petros M. Nomikos Conference Centre in the town of Firá (see photo to the right). The conference will be single-track. Morning talks will be followed by a working lunch. After an afternoon break, participants will reconvene for early evening talks and poster presentations, ending in time for late dinner in accordance with local customs. Although delegates arrive from around the world, AREADNE speakers are selected primarily from the United States, Europe, and the Middle East.

Submission of Poster Abstracts

We anticipate being oversubscribed for 2022 as in previous years, and so strongly encourage anyone who is considering attending the conference to submit a poster abstract. Accepted posters will have one registration reserved for the presenter. Other registration spots will be on a first come first served basis. We are typically able to accommodate only an exceedingly small number of delegates who are not a designated poster presenter. Please see our Call for Abstracts.

Registration and Fees

Window with Flower Conference fees are USD 500 for students, USD 700 for post-docs, USD 900 for faculty members or professionals, and USD 1500 for on-site registrants, space permitting. On-site fees are considered full price; the other levels are discounted thanks to the generosity of our sponsors. Registration fees are payable in USD by credit card. While we will endeavor to keep registration fees as stated here, they are subject to change without notice until registration has opened.

Registration fees cover coffee breaks, hot lunches, the Welcome Reception, the Gala Banquet, and an optional excursion. Attendance will be strictly limited by the capacity of the conference center. Registration is separate from proposal submission. Once registration has filled, a waiting list is established; at previous meetings, only a few people on the waiting list were eventually offered a registration spot. If the conference is full, on-site registrations will not be accepted.

COVID Policy. Detailed pandemic-related restrictions for the meeting will be established once the EU and Greek governmental policies have been set for the conference period. At present, we anticipate that masking and proof of full vaccination including boosters will be required.

Refund Policy. If you have registered but will be unable to attend, a full refund of the paid fees will be given until 31 May 2022, a 50% refund will be given from then until 14 June 2022, and no refund will be given after 14 June 2022.

Registration has closed. We have sold out.

Important Dates

AREADNE 2022 will be Tuesday to Saturday, 28 June - 2 July 2022, starting Tuesday evening with a Welcome Reception. This schedule is a day earlier in the week than previous meetings.

2022-01-03  Abstract submissions open
2022-01-21  Abstract submissions close
2022-02-24  Abstract notifications sent
2022-03-02  Registration opens (subject to change)
2022-06-07  Registration closes
2022-06-27  Suggested arrival date for intercontinental travelers
2022-06-28  Conference Welcome Reception (late PM)
2022-06-29  Conference opens (early AM)
2022-07-02  Conference closes (late PM)
2022-07-02  Conference Gala Banquet (late PM)
2022-07-03  Suggested departure date

Attendees should plan to arrive in Santorini by 28 June 2022 (or the day before, especially for intercontinental travelers), and depart on or after 3 July 2022.


Many hotels are available in and around Firá and Firostefani, from inexpensive C class / two-star hotels to luxury A class / four-star caldera-view cave suites. The conference center is located 15 minutes by foot from the center of Firá, and a similar distance from much of Firostefani. We encourage conference attendees to follow the lead of the organizing committee, and stay in hotels that are walking distance from the Centre. Plan to arrive by Tuesday, 28 June 2022 if coming from Europe or Monday, 27 June 2022 if traveling intercontinentally, and depart Sunday, 3 July 2022.

We suggest booking your hotel room as early as possible.


Blue Urns Daily coffee breaks, lunches, and a Welcome Reception will be provided at the conference center. Working lunches will be provided on three of the four days. A Gala Banquet will be held at a local restaurant on the final day of the conference. These meals are included in the conference package for participants; guests are welcomed to the Welcome Reception, and additional tickets are available for purchase to bring guests to the Gala Banquet (tickets available during on-line registration).

Breakfasts are typically included in hotel rates, or available with bookings for a nominal charge, and are thus not provided at the conference center. There is a wide selection of nearby restaurants for evening meals, and we anticipate making loosely organized excursions to different restaurants each evening, except the last, when we have the banquet.

Restaurant recommendations are available.

Activities on the island

In addition to sweeping vistas, Santorini boasts excellent nightlife, a respectable wine industry, beaches with white, black, or red sand, excavations of ancient civilizations, and magnificent sunsets. More information can be found in various travel guides, or on the Web. Good places to start are publications like The Lonely Planet and The Rough Guide guides for Greece, and web sites like Greek Travel Santorini, Santorini Island, and Travel to Santorini.

We are planning optional guided excursions to the archaeological site of Akrotiri and to Nea Kameni, the volcano island at the center of the caldera. These events may not be able to accommodate everyone.

Additional suggested activities are available.


Church Bell Air transportation to Athens International Airport (ATH) is available through the major international airlines, many discount companies, and numerous charter companies. Air transport from Athens to Santorini (JTR) is available multiple times per day and takes less than an hour. It is usually possible to arrange flight schedules so that overnight stays in Athens are not required when connecting to and from Santorini.

Sea passage from Athens to Santorini can be a relaxing experience and is recommended for the awe-inspiring approach to the port at the base of the immense caldera cliffs. Ferries for Santorini leave from the port of Piraeus, one hour's taxi drive away from the Athens airport, and from the town of Rafina, about 25 minutes by taxi from the airport in the other direction. Rail and express bus transportation are also available from the airport to the ferries in Piraeus.

Getting around on Santorini can be done by bus and taxi. Cars and mopeds are readily available for hire. The conference center is located in the town of Firá, and is walking distance to many hotels and restaurants, so having a motorized vehicle is not required.

More transportation information is available here.

Visa Requirements

European citizens have free Schengen access to Greece. Post-Brexit, UK citizens have visa-free travel to the Schengen zone for 90 days out of 180. United States citizens traveling on regular passports (excluding diplomatic and governmental passports) do not require visas for stays shorter than 90 days. Citizens of other countries will need to contact their home governments to determine visa requirements. We are happy to assist with visa letters for registered attendees.


Bougainvillea with Caldera The schedule for the four-day conference follows the Greek lifestyle of having a long lunch, afternoons free for siestas or swimming, and a late dinner. To encourage participants to stay for the entire event, the detailed speaker and poster schedule will not be available until the start of the conference, even to the speakers and poster presenters. Unlike previous meetings, our schedule for AREADNE 2022 spans Tuesday to Saturday.

All attendees should plan a stay that includes arrival on Santorini no later than Monday, 27 June 2022 for intercontinental travelers or Tuesday, 28 June 2022 for those arriving from Europe and the Middle East, and departure no earlier than the following Sunday, 3 July 2022.

Tuesday, 28 June 2022
19:30-22:00   Welcome Reception (wine and cheese) and Registration

Wednesday, 29 June 2022
08:00-08:30   Registration
08:30-12:30   Lectures and coffee break
12:30-14:00   Lunch
17:00-21:30   Lectures, coffee break, and poster session

Thursday, 30 June 2022
09:00-12:30   Lectures and coffee break
12:30-14:00   Lunch
17:00-21:30   Lectures, coffee break, and poster session

Friday, 1 July 2022
09:00-13:00   Optional archaeological and geological excursions
17:00-21:30   Lectures, coffee break, and poster session

Saturday, 2 July 2022
09:00-12:30   Lectures and coffee break
12:30-14:00   Lunch
17:00-19:30   Lectures and coffee break
21:00-24:00   Gala Banquet (full dinner)

By long-standing policy we do not release the detailed speaker or poster schedule before the start of the conference. (Why not?)

Confirmed Speakers

Our confirmed speakers include many global luminaries in the field of Computational and Systems Neuroscience.

Sliman Bensmaia   University of Chicago
Claudia Clopath   Imperial College
Laura Colgin   University of Texas, Austin
Christine Constantinople   New York University
Ila Fiete   Massachusetts Institute of Technology
Stefano Fusi   Columbia University
Jim Gnadt   National Institude of Neurological Disorders and Stroke
Adam Hantman   Howard Hughes Medical Institute Janelia Research Campus
Elizabeth Hong   California Institute of Technology
Matt Kaufman   University of Chicago
Georg Keller   Friederich Miescher Institute for Biomedical Research
Roozbeh Kiani   New York University
Vassiliki Nikoletopoulou   Institute of Molecular Biology and Biotechnology
Maria Papadopouli   University of Crete
Clay Reid   Allen Institute for Brain Science
John Reynolds   Salk Institute for Biological Studies
Cristina Savin   New York University
Mark Sheffield   University of Chicago
Andreas Tolias   Baylor College of Medicine
Stefan Treue   German Primate Center
Nachum Ulanovsky   Weizman Institute


In addition to our expert speakers, we have an august list of session moderators to keep the discussions lively and interesting.

David Freedman   University of Chicago
Leslie Osborne   Duke University
Barry Richmond   National Institute of Mental Health
Irini Skaliora   BRFAA
Ziv Williams   Massachusetts General Hospital

Organizing Committee

Dora Angelaki   New York University
Kenny Blum   Harvard University
Nicholas Hatsopoulos, co-chair   University of Chicago
John Pezaris, co-chair   Harvard Medical School
Panayiota Poirazi   Foundation for Research and Technology Hellas
Thanos Siapas   Caltech
Andreas Tolias   Baylor College of Medicine

The Myth of Ariadne

Road and Bluff We take the conference name AREADNE from the subtitle, Research in Encoding And Decoding of Neural Ensembles, combined with the name of the mythological figure Ariadne.

Theseus, an Athenian hero, journeyed to Crete to slay the Minotaur, a half-bull, half-human creature. King Minos of Crete kept the Minotaur at the center of a large maze he built, known as the Labyrinth.

Minos' daughter Ariadne, after whom we've named the conference, gave Theseus a sword and a ball of silk thread, both of which had been given to her by Daedalus, designer of the labyrinth. Theseus tied one end of the thread at the entrance, unwinding the rest as he navigated the Labyrinth, so that after slaying the Minotaur, he could easily escape by following the thread back out.

Practical Information

We have assembled a body of practical information to guide planning for attending an AREADNE Conference.

Και τα λοιπά (Et Cetera)

AREADNE 2022 will be Tuesday to Saturday, 28 June - 2 July 2022, starting Tuesday evening with a Welcome Reception. This schedule is a day earlier in the week than previous meetings.

The language of presentation for AREADNE Conferences is English.

To be sent important email announcements about AREADNE 2022, please add yourself to the Google Group for Official AREADNE Announcements.

Internet access will be provided at the Nomikos Conference Centre only to a limited extent. This is a conscious decision to encourage attendees to participate in the conference rather than to be distracted by email and the Web. Internet cafes are readily available in Firá for emergencies and extended access. The response from attendees on this policy was overwhelmingly positive at all of our previous conferences.

Abstracts of all presentations will be published in the conference book that is provided to attendees during registration, and available as a downloadable PDF after the meeting. Copies from previous meetings are available on our archives page.

We have compiled a list of frequently asked questions.

For additional information on other conference-related topics, please contact the Organizing Committee Co-Chairs, Nicholas Hatsopoulos and John Pezaris, at

Photographs on this site were taken on Santorini 2005-2021 by John Pezaris (except the satellite view published by NASA), to display the magnificence and austere beauty of the island. Click on any of them to view a larger version (and use the BACK button on your browser to return to the previous page). Images are copyrighted by the artist and are used by The AREADNE Foundation with permission.

Copyright 2005–2022, All Rights Reserved.
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